Three
types of chaparral occur within the Reserve: the Chamise, Chamise
- Wedgeleaf Ceanothus, and Scrub Interior Live Oak Alliances.
Although these grade into one another, in general the north and
east-facing slopes and ridge tops support the more speciose scrub
oak chaparral. The same areas support chamise - wedgeleaf chaparral
on southerly and westerly trending slopes, and pure chamise is
found on ridgetops and south and west slopes.
Although all of these communities tend to occur in xeric areas,
pure chamise (Adenostoma fasciculatum) claims the driest
areas (12.5%). Chamise has needle-like leaves that are extremely
drought resistant. It is well-adapted to fire by its ability to
resprout very quickly from the root crown as well as to reproduce
prolifically from seed. The area of the Reserve that burned in
1992 shows little sign of disturbance in 2004.
With a canopy comprising 70-80% cover and very little understory,
it is the least diverse community on the Reserve. The lack of
understory plants beneath chamise has been attributed to a variety
of causes over the years, including allelopathic exudates from
the chamise36,
lack of seed sources39,
low moisture and nutrient levels as a result of competition with
chamise roots33,
and herbivory by mammals32,
35, 38.
Many of these factors play a role in the scarcity of understory
cover, but the most compelling seems to be herbivory. The dense
thicket of shrubs provides cover for rabbits and many small rodents
that harvest and eat whatever grasses and forbs manage to germinate
near the canopy.
Scrub oak chaparral (9.2%) contains a mix of several species
in various proportions. As the name implies, two major components
of this community are the shrub form
of
interior live oak (Quercus wizlizeni var. frutescens)
and the scrub oak (Q. berberidifolia). Especially attractive
in the spring, this superalliance also includes flowering ash
(Fraxinus dipetala), toyon (Heteromeles arbutifolia),
California bay (Umbellularia californica), California
buckeye, birchleaf mountain mahogany (Cercocarpus betuloides),
and pitchersage (Lepechinia calycina), all twined with
wild virgin’s bower (Clematis lasiantha). The species
composition appears to be related to aspect and moisture availability.
The
understory of the scrub oak chaparral harbors a mix of small forbs,
lilies, and ferns. Twining brodiaea (Dichelostemma volubile),
Diogenes’ lantern (Calochortus amabilis), shooting
star (Dodecatheon hendersonii), hedge nettle (Stachys
ajugoides), and goldback fern (Pentagramma triangularis)
are some of the natives in this diverse understory.
In many areas scrub oak chaparral interdigitates on the ridges
with the Chamise - Wedgeleaf Ceanothus Alliance, which comprises
7.2% of the Reserve. In general on the northwest-southeast trending
ridges on the west side of the peninsula, the scrub oak occupies
the north and east facing slopes, and the chamise - wedgeleaf
ceanothus the south and west facing slopes. In this community,
as depicted on the map, the co-dominants appear to be a variety
of shrubs including scrub oak, toyon, ceanothus (Ceanothus
oliganthus), birch-leafed mountain mahogany, and interior
liveoak. Wedgeleaf ceanothus (Ceanothus cuneatus) is
uncommon.
Photo Credits: Title, Ribes
malvaceum (Dan Tolson), chamise, California Bay, and Diogenes’
lantern (Ellen Dean). For more pictures see: http://elib.cs.berkeley.edu/photos/flora/